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SVP 2021 abstracts – 10: Apterodon taxonomic issues

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Al Ashqar et al. 2021 bring us their report
on the most complete Apterodon (Fischer 1880; Fig. 1) cranium ever recovered. We looked at Apterodon earlier here (if you want more info on this taxon and its relatives).

Al Ashqar et al. continue:
“Here, we report the most complete Apterodon cranium ever recovered from Quarry M (~29
Ma), one of the youngest terrestrial localities in the Jebel Qatrani Formation of the Fayum Depression. Based on craniodental morphology, the new cranium is very similar to Apterodon macrognathus
[Fig. 1] from Quarry A (~ 33 Ma), also in the Jebel Qatrani Formation.”

“The new skull preserves the complete upper left tooth row aside from I1 and P1 and right I2, P2-3 and M2. Like other apterodontines, the carnassial cutting blade is very reduced, reflecting a possible mesocarnivorous diet.”

That’s not a carnassial in Apterodon, it’s a molar. Purported sisters described by Al Ashqar are not related to one another in the LRT. He’s following a tradition based on taxon exclusion.

Al Ashqar et al. continue:
The skull is anteroposteriorly elongate with a narrow, crushed rostrum. The nuchal crest and basicranium are well preserved with two large occipitals. The nuchal crest has a clover shape and does not reach the mastoid process. The neurocranium is complete and elongated. The zygomatic arches are dorsoventrally tall, broad and robust. The petrosal is preserved in the right side. The discovery of this cranium allows detailed craniodental comparisons with the better-known A. macrognathus, better documentation of the evolution of the distinct
apterodontine bauplan, and closer examination of the transtethyan dispersal of Apterodon during the Paleogene.”

Also added to the LRT is the related Pterodon, aka Hyainailouros (Fig. 3) which also nests with Anagale (Fig. 4). If only Al Ashqar had mentioned that they also studied anagalids, then we’d have something to talk about. Sometimes it’s better to look at other taxa (= comparative anatomy) rather than looking more closely at the specimen under review.

Anagalids spread worldwide throughout the Cretaceous. Anagale is a late survivor form the Early Oligocene. The Early Eocene IVPP V12385 specimen assigned to Hapalodectes (Fig. 5) is also a taxon close to Apterodon.


Figure 5. The IVPP V 12385 specimen of Hapalodectes compared to the holotype specimen. The former nests with Anagale. The latter nests with Ptilocercus.

Once again, adding taxa clarifies interrelationships. With 1990 taxa, the LRT is able to lump and separate taxa that are traditionally confused with one another due to taxon exclusion.

References
Al Ashqar et al. 2021. The cranium of Apterodon (Hayainalouridae, Apterodontidate) from one of the youngest vertebrate bearing locatlies (Early Oligocene) in the Fayum Depression, Eqypt. SVP abstracts 2021: 44.
Fischer P 1880. Note sur un nouveau genre de mammifere fossile (Apterodon gaudryi) des Phosphorites du Quercy. Bulletin de la Société Géologique de France 8: 288-290.

wiki/Apterodon
wiki/Hyaenodon
wiki/Hyainailouros
wiki/Anagale


Source: https://pterosaurheresies.wordpress.com/2021/11/09/svp-2021-abstracts-10-apterodon-taxonomic-issues/


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