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Glyptolepis, a wide-skull lobe-fin fish, enters the LRT and splits up the Porolepiformes

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Glyptolepis paucidens (Agassiz 1844; Middle Devonian; 20cm; Figs. 1, 2) was a smallish rhipidistian, a lobefin fish with primitive lobes. Traditionally considered a porolepiform, after testing Glyptolepis does not nest with the porolepiform, Porolepis (Fig. 3), which nests several nodes apart in the LRT, closer to coelocanths.

Here
in
the large reptile tree (LRT, 2021 taxa) Glyptolepis nests closer to the tetrapodomorph, Cabonnichthys (Fig. 4), far from Porolepis and Holoptychius (Fig. 3), despite several convergent traits. The porolepiform, Laccognathus (Fig. 5), has a flatter, wider skull similar to that of Glyptolepis by convergence. Moving Glyptolepis to Porolepis or vice versa adds 30 steps to the LRT.

Porolepis posnaniensis
(Kade 1858, named by Woodward 1891; Early Devonian) was a large, slow-moving lobefin with small eyes, related to Holoptychius, not Glyptolepis, contra academic traditions.

Ahlberg 1991
re-examined Porolepiformes at the start of the software era of cladistics. Ahlberg reported,
“A cladistic analysis based on 54 characters places the Porolepiformes as sister-group of a clade
Powichthys + Youngolepis + Diabolepis + Dipnoi. The porolepiform-lungfish clade is the sister group of another clade containing tetrapods, panderichthyids, osteolepiforms and rhizodonts.”

The LRT employs more taxa.
The LRT widely separates porolepiforms from lungfish, which are the sister clade to tetrapodomorphs llike panderichthys, osteolepiforms and rhizodonts. The LRT nests Youngolepis closer to Porolepis than to lungfish (= Dipnoi). These relationships await confirmation with a similar taxon list.

This appears to be a novel hypothesis of interrelationships.
If not, please provide a citation so I can promote it here. This discovery was accomplished by simply adding taxa, letting the software decide interrelationships and not assuming that similar traits indicate relationships rather than convergence. We’ve seen this before with
‘turtle’ origins, ‘whale’ origins, pterosaur origins and more.

References
Ahlberg PE 1991. A re-examination of sarcopterygian interrelationships, with special reference to Porolepiformes. Zoological Journal of the Linnean Society 103(3):241–287,
Ahlberg PE and Johanson Z 1997. Second tristichopterid (Sarcopterygii, Osteolepidformes) from the Upper Devonian of Canowindra, New South Wales, Australia, and phylogeny of the Tristichopteridae. Journal of Vertebrate Paleontology 17(4):653–673.
Cloutier R and Ahlberg PE 1996. Morphology, Characters, and the Interrelationships of Basal Sarcopterygians. In Melanie LJ Stiassny; LR Parenti and  GD Johnson (eds.). Interrelationships of Fishes. p. 457.
Kade G 1858. Ueber die devonischen Fischreste cines diluvial Blockes. Meseritz. pp.17-18. Woodward AS 1891. Catalogue of the Fossil Fishes in the British Museum (Natural History). Part II. Catalogue of the Fossil Fishes in the British Museum (Natural History) 2.

wiki/Glyptolepis
wiki/Cabonnichthys
wiki/Porolepis
wiki/Porolepiformes


Source: https://pterosaurheresies.wordpress.com/2021/12/25/glyptolepis-a-wide-skull-lobe-fin-fish-enters-the-lrt-and-splits-up-the-porolepiformes/


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