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Early Eocene Micrornatus enters the LRT, but not with Scomber, the mackerel

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From the Beckett and Friedman 2015 abstract:
“The monotypic scombrid fish Micrornatus is represented by a single skull from the
early Eocene (Ypresian) London Clay Formation of southeastern England. Here we re-examine this specimen using computed microtomography. Scans reveal new details of the braincase, suspensorium and ventral hyoid arch. “

The huge (35cm) skull of Micrornatus is shown in figure 1.

It’s good to see workers coloring bones, but some standardization would be better. Here the authors’ colors are replaced with tetrapod homology colors used at ReptileEvolution.com (e.g. premaxilla = yellow, maxilla = green, etc). You might notice a few bones here (e.g. the nasals and frontals, the dentary and articular) are reidentified.

Micrornatus hopwoodi
(Casier 1966, formerly Eocoelopoma hopwoodi, Beckett and Friedman 2015, early Eocene, BMNH 36136 = NHMUK PV OR 36136) was considered a mackerel close to Scomber, but in
the large reptile tree (LRT, 2206 taxa) nests with Esox, the muskellunge (muskie, Fig 2) a type of pike. The premaxilla and its teeth on marine Micrornatus are much larger than on the freshwater pike, which has larger dentary teeth.

Micrornatus is more muskie (Esox) than mackerel (Scomber).
This is what the LRT recovers (Fig 3). Micrornatus was likely a late survivor in the early Eocene.


Figure 3. Recently updated subset of the LRT focusing on Scomber its relatives along with more distantly related taxa that might be confused with Scomber (e.g. Scomberoides, Scomberomorous).

TAt this point, eleven years into the project,
the LRT (subset Fig 3) needs confirmation, refutation or modification by workers employing the same taxon list and a list of their own 200+ random characters.

References
Beckett HT and Friedman M 2015. The one that got away from Smith Woodward: cranial anatomy of Micrornatus (Acanthomorpha: Scombridae) revealed using computed microtomography. In: Johanson Z Barrett PM, Richter M and Smith M (eds) Arthur Smith Woodward: His Life and Influence on Modern Vertebrate Palaeontology. Geological Society, London, Special Publications, 430, http://doi.org/10.1144/SP430.16
Pauca M 1929. Vorläufige Mitteilung über eine fossile Fischfauna aus den Oligozänschiefern von Suclânesti (Muscel). Acad. Roum. Sect. Sci. Bull. v. 12, p. 112-121.

wiki/Macrornatus – not yet posted


Source: https://pterosaurheresies.wordpress.com/2023/01/29/early-eocene-micrornatus-enters-the-lrt-but-not-with-scomber-the-mackerel/


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