The preoperculum and the ‘preoperculum’ are not homologous
According to Wikipedia,
“The preoperculum is a crescent-shaped structure that has a series of ridges directed posterodorsally to the organisms canal pores. The preoperculum can be located through an exposed condyle that is present immediately under its ventral margin; it also borders the operculum, suboperculum, and interoperculum posteriorly.”
The wiki-authors are referring to the majority
of preoperculum appearances in vertebrates (Fig.1 ), ignoring the others (Figs. 2-3).
A preoperculum can also appear
as a bone posteriorly bordering the squamosal and quadratojugal (Figs. 2-4). Intervening taxa have no preoperculum, so the various appearances of this bones are not homologous, but keep returning as reversals. Note the appearance of a preoperculum in most extant fish and in unrelated catfish, too.
Among tetrapods
the preoperculum makes a brief reappearance in the Ossinodus (Fig. 4) clade.
Source: https://pterosaurheresies.wordpress.com/2020/02/23/the-preoperculum-and-the-preoperculum-are-not-homologous/
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