Subadult and adult Tropeognathus compared
Holgado and Pegas 2020 name several toothy crested rostral pieces
they assign to Anhangueridae, Coloborhynchinae and Tropeognathinae, subsets of the clade Ornithocheiridae (Seeley 1870).
More interesting due to its completeness,
was the Holgado and Pegas page-wide photo of the BSp 1987 I 47 specimen of Tropeognathus (Fig. 1, Wellnhofer 1987). Here the ’47’ specimen is compared to the larger holotype, BSp 1987 I 46, and to the smaller Scaphognathus holotype (Fig. 1, GPIB 1304, No. 109 of Wellnhofer 1975), a distant ancestor of Tropeognathus in the Large Pterosaur Tree (LPT, 251 taxa).
Holgado and Pegas provided a cladogram of a clade of pterosaurs
formerly considered Ornithocheiridae, Lanceodontia. Strangely, their outgroup is the derived istiodactylid, Lonchodraco giganteus, which we looked at earlier here and nested with the unnamed SMNS PAL 1136 specimen, which was omitted from their cladogram.
By contrast
In the LPT ornithocheirids arise from small taxa like Yixianopterus, Mimodactylus and before them the Cycnorhamphus clade and before them the tiny Late Jurassic pterosaurs, BM NHM 42735, Gmu10157, TM 13104 and three Scaphognathus specimens (109, SMNS 59395 and 110 and of descending size. None of these are included in Holgado and Pegas.
There are quite a few nomenclature problems
in the Ornithocheridae that make the taxonomy unnecessarily confusing.
According to Wikipedia,
“Back in 1987, Wellnhofer had named a second species called Tropeognathus robustus, based on specimen BSP 1987 I 47, which is a more robust lower jaw. In 2013 however, T. robustus was considered as a species of Anhanguera, resulting in an Anhanguera robustus.” Comparing the 46 and 47 specimens (Fig. 1) show they are conspecific, or at least congeneric. This clade becomes increasingly confused with every new author or set of authors. Strange that such closely related taxa are generically split while the several dozen variations in Rhamphorhynchus and Pteranodon are ignored.
The first species of Tropeognathus mesembrinus has several synonyms.
- Anhanguera mesembrinus (Wellnhofer, 1987)
- Coloborhynchus mesembrinus (Wellnhofer, 1987)
- Criorhynchus mesembrinus (Wellnhofer, 1987)
- Ornithocheirus mesembrinus (Wellnhofer, 1987)
Ornithocheiridae Seeley 1870, named when only a few bits and pieces were known
Ornithocheiromorpha Andres et al., 2014, incorrectly nested within Pteranodontoidea.
Pterodactyloidea Plieninger, 1901, adding taxa splits up this traditional clade.
The subadult specimen (specimen ’47’)
of Tropeognathus (Fig. 1) documents a vestige, slit-like naris that disappears in the larger ’46’ specimen. If you can’t see it here, click to enlarge.
Rostral crest
Comparing the subadult to the adult specimen (Fig. 1) demonstrates no growth in the size of the rostral crest. Rather the back half of the skull is slightly larger.
References
Holgado B and Pegas RV 2020. A taxonomic and phylogenetic review of the anhanguerid pterosaur group Coloborhynchinae and the new clade Tropeognathinae. Acta Palaeontologica Polonica 65 (X): xxx–xxx.
Wellnhofer P 1975a. Teil I. Die Rhamphorhynchoidea (Pterosauria) der Oberjura-Plattenkalke Süddeutschlands. Allgemeine Skelettmorphologie. Paleontographica A 148: 1-33. 1975b. Teil II. Systematische Beschreibung. Paleontographica A 148: 132-186. 1975c.Teil III. Paläokolgie und Stammesgeschichte. Palaeontographica 149: 1-30.
Wellnhofer P 1987. New crested pterosaurs from the Lower Cretaceous of Brazil. Mitteilungen der Bayerische Staatssammlung für Paläontologie und historische Geologie 27: 175–186; Muenchen
/wiki/Tropeognathus
wiki/Scaphognathus
wiki/Criorhynchus
Source: https://pterosaurheresies.wordpress.com/2020/09/27/subadult-and-adult-tropeognathus-compared/
Anyone can join.
Anyone can contribute.
Anyone can become informed about their world.
"United We Stand" Click Here To Create Your Personal Citizen Journalist Account Today, Be Sure To Invite Your Friends.
Please Help Support BeforeitsNews by trying our Natural Health Products below!
Order by Phone at 888-809-8385 or online at https://mitocopper.com M - F 9am to 5pm EST
Order by Phone at 866-388-7003 or online at https://www.herbanomic.com M - F 9am to 5pm EST
Order by Phone at 866-388-7003 or online at https://www.herbanomics.com M - F 9am to 5pm EST
Humic & Fulvic Trace Minerals Complex - Nature's most important supplement! Vivid Dreams again!
HNEX HydroNano EXtracellular Water - Improve immune system health and reduce inflammation.
Ultimate Clinical Potency Curcumin - Natural pain relief, reduce inflammation and so much more.
MitoCopper - Bioavailable Copper destroys pathogens and gives you more energy. (See Blood Video)
Oxy Powder - Natural Colon Cleanser! Cleans out toxic buildup with oxygen!
Nascent Iodine - Promotes detoxification, mental focus and thyroid health.
Smart Meter Cover - Reduces Smart Meter radiation by 96%! (See Video).